Fig. 4. SMC2 is required for proper meiotic chromosome segregation. Pollen mother cells at both metaphase I and anaphase I from wild-type (A,D) and E1^+/-E2^-/- plants (B,C,E,F) were stained with DAPI and examined by fluorescence microscopy. In wild type, discrete signals and proper segregation of chromosomes were observed, whereas in a population of the mutant pollen mother cells, either relatively normal (B,E) or diffuse signals and chromatin bridges were often observed (C,F).