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Fig. 3. The role of TSH in disco mRNA distribution. (A) disco mRNA accumulation in a wild-type embryo as the germband begins to retract. Segment abbreviations are as in Fig. 1. Note the spots of expression in the thorax and abdomen, indicated by white arrows. In the thorax, these demark the Keilin's Organ precursors. (B) In homozygous tsh8 embryos, gnathal expression of disco is normal, but the trunk distribution is altered. There is a wider distribution in the ventral and ventral-lateral region of trunk segments, particularly notable in the thoracic segments. Note that the spots marking the Keilin's Organs are missing. (C) Ectopic activation of disco caused by Gal4-driven DFD. Interestingly, the distribution of disco mRNA is quite similar to that in B, above, except that the spots marking the Keilin's Organ precursors are still present in C. (D-F) Ectopic expression of tsh represses the normal accumulation of disco mRNA in the gnathal segments. (D) Wild-type late stage 12 embryo. disco mRNA distribution is fairly uniform in the gnathal segments except where the maxillary and labial sense organs will form and in the salivary primordia (ventral labial). (E) In arm->tsh embryos (stage 12), as the germband retracts, disco mRNA diminishes in the epidermis of the gnathal lobes. However, staining increases in the central region of the mandibular, maxillary and labial lobes. Later (stage 13) (F), the labial lobe has taken on the appearance of a first thoracic segment, and disco is strongly expressed in the sensory precursor, which has been transformed toward a Keilin's Organ. No difference was noted in the trunk disco expression.