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Fig. 7. Delta1 expression is upregulated by Xiro1 and down regulated by Snail. (A,B) Two-cell embryos were injected in one blastomere with 1 ng HDGREnR (A) or with 1 ng HDGR (B) mRNA. The embryos were treated with dexomethasone at stage 12 and the expression of Delta1 was analyzed by in situ hybridization at stage 17. The injected side was visualized by alkaline phosphatase-mediated FITC immunodetection. (A) HDGREnR produces a moderate expansion of Delta1 expression in the neural crest region (arrowhead), whereas HDGR leads to a complete inhibition of Delta1 expression in the crest region (B; arrowhead). (C-E) Animal caps taken from stage 9 embryos were cultured until the equivalent of stage 18, and the expression of Delta1 was analyzed by in situ hybridization. (C) In control animal caps, no expression of Delta1 could be detected. (D,E) In animal caps taken from embryos injected with 1 ng Xiro3 mRNA, Delta1 expression was observed in 87% of the caps (n=57). (E) Higher magnification of the animal cap shown in D. (F) RT-PCR to analyze Delta1 and H4 mRNA. Arrowheads in D,E indicate Delta1-expressing cells. Left panel, control embryo and PCR in the absence of reverse transcriptase; right panel, mRNA taken from a control animal cap, a cap injected with 1 ng Xiro3 mRNA, or a cap co-injected with 1 ng Xiro3 mRNA and 0.7 ng Xsnail mRNA. (G) Quantification of data shown above in F. Note the increase in Delta1 mRNA produced by Xiro3, and the complete inhibition produced by Xsnail. (H-J) Two-cell embryos were injected in one blastomere with 0.7 ng dnSnail mRNA. The embryos were treated with dexomethasone at stage 12, and the expression of Delta1 was analyzed by in situ hybridization at stage 17. The injected side was visualized by alkaline phosphatase-mediated FITC inmunodetection. (H) Delta1 expression is upregulated in the neural crest region. (I,J) Higher magnification of the neural crest region indicated by the box in H, where the staining was stronger on the injected (J) than on the uninjected side (I). Arrow, uninjected side; arrowhead, injected side. Each experiment was performed at least twice with a minimum of 52 embryos. The effect seen in each experiment was observed in at least 65% of embryos.