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Fig. 4. Proposed model of the functional complexes of C. elegans Smads. The body size control requires a SMA-3:SMA-4:SMA-2 complex and the feedback loop needs a SMA-2:SMA-4:SMA-3 complex; subunits are shown in an A,B,C configuration, as defined by Chacko et al. (Chacko et al., 2004).