Fig. 7. Phenotypes of double mutants between roxy1-3 and ap2-5, lug-1, ufo-2 and rbe-2. (A-D) Enhancement of first whorl carpeloidy of roxy1 in combination with negative regulators of AG. (A) ap2-5 mutant flower showing transformation of sepals into carpeloid organs. (B) In roxy1-3 ap2-5 double mutants, first whorl carpeloidy is strongly enhanced and fused organs are topped with stigmatic papillae. (C) The lug-1 single mutant forms feminized first whorl organs. (D) In the roxy1-3 lug-1 double mutant carpeloidy of the first whorl organs is enhanced. (E) Mutants of the strong ufo-2 allele display defects in all floral whorls, affecting most severely second and third whorl organs resembling sepaloid and stamenoid structures, respectively. (F) roxy1-3 ufo-2 double mutant flowers form a reduced number of carpeloid second whorl organs. One first whorl sepal was removed to reveal inner organs. (G) rbe-2 single mutant, where sepals were removed. Arrows indicate filamentous structures formed instead of petals in the second whorl. (H) roxy1-3 rbe-2 double mutant flower, sepals were removed. Extra stamens develop in the second whorl (arrowheads), instead of petals or filamentous structures. ls, lateral stamen.