Cover: Mouse E16.5 dorsal tongue filiform papillary epithelia are highly patterned as rosettes, ordered between regions of intercalating lamina propria. Epithelium is demarcated by membranously expressed E-cadherin (Rhodamine Red-X); nuclei are stained with DAPI (blue). The mitotic spindle orientation factor LGN has multiple distinct functions in regulating oral epithelial development by controlling cell division orientation. See Research article by Byrd et al. on p. 2803.